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    Robert Guthrie (microbiologist)

    American microbiologist (–)

    Robert Guthrie, MD, Ph.D. (June 28, &#; June 24, ) was an American microbiologist, best known for developing the bacterial inhibition assay used to screen infants for phenylketonuria at birth, before the development of irreversible neurological damage.[1] Guthrie also pioneered the collection of whole blood on specially designed filter paper, commonly known as "Guthrie cards" as a sample medium that could be easily collected, transported and tested.[2] Although Guthrie is best known for developing the test for phenylketonuria, he worked tirelessly to raise awareness of the need to screen for treatable conditions and adapted his method to early screening tests for galactosemia and maple syrup urine disease.

    Early life

    Guthrie received his doctorate from the University of Minnesota,[3] although his education took a circuitous route, as he eventually earned six degrees in six years, inc